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Population Genetics of Parasites and Their Arthropod Hosts
Bezányiová, Kateřina ; Straka, Jakub (advisor) ; Votýpka, Jan (referee)
Arthropoda are currently the largest metazoan phylum. Given that organisms with parasitic lifestyle are thought to comprise the majority of existing species, it's easy to imagine an immense diversity of parasites interacts with arthropods. However, in comparison to organisms parasitising vertebrates, parasites of arthropods are direly understudied despite their abundance, importance, and potential usefulness. Amongst other things, parasites can be used as tools allowing the inference of information on host life history, ecology, and past events the host species have experienced. Population genetic structure of parasites and other symbionts may reflect these traits and events due to their close relationship with the host. Even though parasites comprise a diverse assemblage of taxa, it's possible to identify convergent patterns in their biology. Models predicting congruent population genetic co- structuring can be thus based on a few traits such as host specificity, life cycle complexity or parasite and/or host dispersal. In some cases, the parasite may provide better resolution of population structure than the host itself, serving as a proxy that may be used to direct conservation programmes of both the host and parasite, as has already been done with parasites of vertebrates. This thesis summarises known...
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Comparison of Genetic Programming Variants in the Symbolic Regression Task
Doležal, Petr ; Hurta, Martin (referee) ; Drahošová, Michaela (advisor)
This thesis deals with comparison of genetic programming variants it the task of symbolic regression. Time to converge and quality of evolved solutions are evaluated on nine chosen benchmarks. In particular, tree-based genetic programming, cartesian genetic programming and their modifications using coevolutionary algorithm are investigated. An own implementation of employed methods (without a specific library use) allows to share as much code as possible. Moreover, an analysis of implemented methods efficiency on real world data is provided. Experimental results show that all of the investigated approaches are capable of finding solutions using symbolic regression. Cartesian genetic programming enhanced with coevolution seems to be the most suitable of the investigated approaches in terms of evolved solution quality and time to converge.
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Coevolution of AI and level generation for Super Mario game
Flimmel, Július ; Černý, Vojtěch (advisor) ; Pilát, Martin (referee)
Procedural Content Generation is now used in many games to generate a wide variety of content. It often uses players controlled by Artificial Intelligence for its evaluation. PCG content can also be used when training AI players to achieve better generalization. In both of these fields, evolutionary algorithms are employed, but they are rarely used together. In this thesis, we use the coevolution of AI players and level generators for platformer game Super Mario. Coevolution's benefit is, that the AI players are evaluated by adapting level generators, and vice versa, level generators are evaluated by adapting AI players. This approach has two results. The first one is a creation of multiple level generators, each generating levels of gradually increased difficulty. Levels generated using a sequence of these generators also mirror the learning curve of the AI player. This can be useful also for human players playing the game for the first time. The second result is an AI player, which was evolved on gradually more difficult levels. Making it learn progressively may yield better results. Using the coevolution also doesn't require any training data set.
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Animal phobia: fear or disgust?
Peléšková, Šárka ; Landová, Eva (advisor) ; Hocko Fajnerová, Iveta (referee)
Animal phobias are among the most common specific phobias. It is traditionally assumed that two basic emotions are involved - fear and anxiety, although recent studies suggest that disgust may also be important. Phobias and involved emotions are defined mostly by psychologists and psychiatrists. Their manifestations could be identified mainly on physiological level and are being researched using advanced medical methods (e.g. functional neuroimaging). The eliciting stimuli of arachnophobia, ophidiophobia etc. are specific animal, spiders or snakes. However the category of elicitors for disgust sensu lato is fairly extended. The etiology of these phenomena is explained from the perspective of evolutionary biology and presumes biologicaly significant selective pressures and common coevolution. The aim of this paper is to summarize information gathered so far, verify proposed evolutionary hypothesis about the involvement of disgust in the etiology of animal phobia and to evaluate biological relevancy of fear of animal stimuli.
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Phylogeny of selected species groups of the genus Torymus (Hymenoptera: Torymidae)
Křížková, Barbora ; Janšta, Petr (advisor) ; Skuhrovec, Jiří (referee)
Chalcid wasps (Chalciodiodea) is superfamily within parasitic group of the order Hymenoptera. These insects attack other arthropods. My thesis is focused on phylogeny of selected species groups of the genus Torymus Dalman 1820, whose range of hosts includes mainly gall-forming insect as gall wasps (Hymenoptera: Cynipidae) and gall midges (Diptera: Cecidomyiidae). Using genes 28D2 rDNA, COI and CytB, I constructed phylogenetic trees, which helped with discussing current view on classification of the genus and with inferring new findings about co-evolution with host organisms. Previously established morphological - ecological species groups were not supported by my analyses in many cases. Morphological traits seem to be convergent for many species. They often do not support observed monophyla as apomorphies. On the other hand, ecology and natural habitat of species was common for many branches in my cladograms. It was found out, that recent hosts are not probably original hosts of the group. Adaptive radiation in different habitats is likely to be the phenomenon that stands behind present-day host range of Torymus species. This process could have occurred after swapping to gall midges and gall wasps. Key words: Chalcidoidea, Torymus, phylogeny, parasitoid, host specifity, co-evolution
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Preferences of the main groups of pollinators for the functional floral traits
Malá, Karolína ; Janovský, Zdeněk (advisor) ; Hadrava, Jiří (referee)
This bachelor thesis summarizes the preferences of the main functional groups of pollinators towards floral traits of plants. Due to coevolution and mutual selection pressures, these two life forms have begun to adapt to each other and develop traits on which the future of their life depends. The plant seeks to attract its pollinators, thus ensuring efficient pollen transfer or reproduction for its species. Attractants are used to motivate visitors to its flowers. These attractants provide pollinators with either direct rewards (food, shelter, brood) or rewards in the form of sensory perceptions (colour, scent, morphology). Recently, it has become apparent that flower colour, morphology and the type of food reward offered by the flower have the greatest influence on pollinator choice. Based on preferences towards attractants, we can organize several functional groups whose representatives have the same or similar choice preferences. Key words Pollination syndromes, pollinator, selection pressures, plant traits, coevolution, preferences, fidelity, diptera, coleoptera, lepidoptera, hymenoptera, birds, colour, morphology, rewards
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Phylogeny of selected species groups of the genus Torymus (Hymenoptera: Torymidae)
Křížková, Barbora
Chalcid wasps (Chalciodiodea) is superfamily within parasitic group of the order Hymenoptera. These insects attack other arthropods. My thesis is focused on phylogeny of selected species groups of the genus Torymus Dalman 1820, whose range of hosts includes mainly gall-forming insect as gall wasps (Hymenoptera: Cynipidae) and gall midges (Diptera: Cecidomyiidae). Using genes 28D2 rDNA, COI and CytB, I constructed phylogenetic trees, which helped with discussing current view on classification of the genus and with inferring new findings about co-evolution with host organisms. Previously established morphological - ecological species groups were not supported by my analyses in many cases. Morphological traits seem to be convergent for many species. They often do not support observed monophyla as apomorphies. On the other hand, ecology and natural habitat of species was common for many branches in my cladograms. It was found out, that recent hosts are not probably original hosts of the group. Adaptive radiation in different habitats is likely to be the phenomenon that stands behind present-day host range of Torymus species. This process could have occurred after swapping to gall midges and gall wasps. Key words: Chalcidoidea, Torymus, phylogeny, parasitoid, host specifity, co-evolution
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Coevolution of AI and level generation for Super Mario game
Flimmel, Július ; Černý, Vojtěch (advisor) ; Pilát, Martin (referee)
Procedural Content Generation is now used in many games to generate a wide variety of content. It often uses players controlled by Artificial Intelligence for its evaluation. PCG content can also be used when training AI players to achieve better generalization. In both of these fields, evolutionary algorithms are employed, but they are rarely used together. In this thesis, we use the coevolution of AI players and level generators for platformer game Super Mario. Coevolution's benefit is, that the AI players are evaluated by adapting level generators, and vice versa, level generators are evaluated by adapting AI players. This approach has two results. The first one is a creation of multiple level generators, each generating levels of gradually increased difficulty. Levels generated using a sequence of these generators also mirror the learning curve of the AI player. This can be useful also for human players playing the game for the first time. The second result is an AI player, which was evolved on gradually more difficult levels. Making it learn progressively may yield better results. Using the coevolution also doesn't require any training data set.
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Host specificity, genetic variability and genealogy in populations of model parasite species
MARTINŮ, Jana
Parasites represent one of the most common ecological strategies and host-parasite coevolution belongs among the major processes governing evolution of biodiversity on the global scale. Genetic structure and diversity of populations of parasites and their hosts, and their genetic connectivity are the key elements in long-term population survival and evolution. Host switches often disturb the parallel evolution of interacting taxa, even in highly host-specific parasites. Evaluation of importance of the degree of intimacy between parasites and hosts is not a trivial task, because evolutionary patterns observed today were formed by an interplay of many (sometimes previously unforeseen) historical and ecological factors. To reveal the mechanisms of coevolution between parasites and their hosts, inter- and intra-specific genealogical structures in three model systems were analyzed: namely, the sucking lice Polyplax serrata and Apodemus hosts, chewing lice of the genus Menacanthus and endoparasitic coccidean genus Eimeria from Apodemus mice.
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Coevolution of avian ectoparasites in the tropics
Gajdošová, Magdalena ; Munclinger, Pavel (advisor) ; Votýpka, Jan (referee)
Host-parasite associations are born by cospeciations or by host switches. Feather lice have traditionally been used as model parasites for studying these events and underlying ecological factors. By now tens of analyses have addressed comparisons of host and parasite phylogenies to study cospeciations and host switches in lice, however, these analyses are strongly biased towards the temperate zone. Tropical environment could provide new insight into the origin of host-parasite interactions, because it is ecologically unique. This work aims to supplement the knowledge of host-parasite associations in lice using coevolution analyses of two feather lice genera and their passerine hosts in tropical rainforest in Cameroon. It shows that lice in the tropics cospeciate rarely. To assess whether host switches are non-random and occur preferentially between hosts with specific traits, this work also analyses relations between parasite genetic distances and hosts' trait similarities. No effect of host morphology and spatial distribution was found. However, genetic distances of the lice strongly correlate with genetic distances of their hosts.
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